A No. 1 Mk. III mag has two locking lugs on the back. A No. 4's has one. They are not interchangeable as issued.
No. 1 mags can be made to fit a No. 4 by grinding the lower lug off. Kind of expensive thing thing to do these days.
A No. 5 Rifle is just a modified No. 4. Same mag.
Like I said, you can and I have used a No4 magazine in a No1Mk3, but you can't use a no1 in a no4/5. They are not REVERSE interchangeable.