The difference is the type of crimp. The 'factory crimp' puts a squeeze from outside the case all the way around the bullet. This will also put a cannelure onto plain bullet if it doesn't have one. It also will NEVER buckle a case.
The roll crimp is just that; it 'rolls' the lip of the case neck into a cannelure on a bullet. Seat the die in too far and you will buckle the neck (or the case if you're loading pistol ammo), and give you all kinds of grief when you try to load/chamber the round. ALL Lee bullet seating dies will roll crimp; read the instructions and they tell you how. The same die, set differently, only seats the bullet and lets you use the Factory Crimp Die. You cannot roll crimp a jacketed bullet without a cannelure; that's the biggest reason people go with the factory crimp; it's simple, works with ANY bullet, and makes seating depth a little less critical because it provides a consistent starting pressure to get the bullet moving.