A sack of bolt heads helps. Armourers had access to a large supply and could pick and choose. Trial and error.
If you stop and think about it, the case rim sits against the barrel face. When the bolt is closed,, there is a gap between the bolt face and the barrel face. This gap should be at least .064", and no more than .074". Less is better. That is why the military gauges are .064" and .074". Case rims should be at least .060". If they are thinner (which happens), headspace will be on the loose side, even if the rifle gauges within limits.
And, of course, bolt heads must turn in just past lining up with the bolt rib.
Your '43 accepts the .074" gauge. If you handload for it, this is irrelevant. Treat the cases as if rimless, and headspace on the shoulder. Never set the shoulder back any more than necessary for easy chambering. First firing of factory ammunition can be done with little rubber bands slipped onto the case. These will hold the rims back against the bolt face, and the cases will fireform.
Lee Enfields can be quirky.