Leading can be caused by too soft an alloy for a given velocity, too hard an alloy for a given velocity, undersized bullets, too high a velocity for the twist rate, too high a velocity for the lube type, too low a velocity for the lube type, and many other things. It is commonly caused by undersized bullets but I find it most commonly caused by too high a velocity for a given hardness and twist rate (too high an RPM) in my own shooting.
I've heard of CZ's slugging odd sizes; both smaller and larger than normal depending on the age/model of the gun. Also see if you can slug just the throat and see if it's significantly different.