what's the diff?

short answer - nope.

The way they fit in the receiver socket is different, plus the shape / size of the rear side is different (different butt plate).

Lou
 
There was no fitting required when I put this Longbranch butt on a 1916 BSA Mklll bitster sniper wannabe.

EnFul2.jpg
 
The safety cutout on the No. I is different. There are at least two patterns of butt socket recess used in No. 4s. As mentionned, adaptation is possible, but the stocks are not directly interchangable.
 
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