I wouldn't use the Fiochhi or Hornady ammo in the Mk.I. The Mk.I was made from a lower grade of steel than later marks, and if they haven't been upgraded, they lack the hardened, replaceable recoil shield too. For that matter, my first Mk.VI came with four boxes of the Fiochhi ammo, and the revolver loosened up quite noticeably by the time that ammo was gone. It seems a bit too snappy for older revolvers.
Many of the Webleys have been fired quite a bit with .45ACP ammo after being shaved, but I think the undersize bullets lower the pressure quite a bit. If you loaded to ACP levels with properly fitting bullets, you'd likely blow the gun up. There is really no point in trying to soup up the .455, in any case. A target really doesn't care if it's hit by a bullet going 550fps, or 800.